In this section we consider small populations (i.e. for which ) so that as a first approximation one can neglect environmental but not demographic stochasticity. Just because populations are very small, one can often assume that the rate of demographic growth does not depend on density and dynamics is therefore Malthusian. To facilitate understanding of the main concepts that guide the analysis, it is convenient to assume population dynamics in continuous rather than discrete time. However, the results would not change qualitatively if one considered the mathematically more complex case of discrete reproduction. Recall that the instantaneous birthrate and death-rate can be reinterpreted in terms of individual fitness as
probability that a female produces a daughter in the small time interval
probability for a female to die in the small time interval
The analytical approach to demographic stochasticity is not easy. We simply summarize the most important results in the Malthusian case ( = constant, = constant, both independent of ) and then briefly treat the case with density dependence. The reader can refer to Iannelli and Pugliese (2014) for a more thorough analysis.